@import url('https://fonts.googleapis.com/css2?family=Inter:wght@400;500;600&display=swap');

:root {
  --canvas: #f7f7f4;
  --canvas-soft: #fafaf7;
  --surface-card: #ffffff;
  --surface-strong: #e6e5e0;
  --hairline: #e6e5e0;
  --hairline-soft: #efeee8;
  --hairline-strong: #cfcdc4;
  --ink: #26251e;
  --body: #5a5852;
  --body-strong: #26251e;
  --muted: #807d72;
  --muted-soft: #a09c92;
  --primary: #f54e00;
  --primary-active: #d04200;
  --on-primary: #ffffff;
  --success: #1f8a65;
  --error: #cf2d56;
}

*, *::before, *::after {
  box-sizing: border-box;
  margin: 0;
  padding: 0;
}

html {
  scroll-behavior: smooth;
}

body {
  background-color: var(--canvas);
  color: var(--body);
  font-family: 'Inter', system-ui, 'Helvetica Neue', Helvetica, Arial, sans-serif;
  font-size: 16px;
  line-height: 1.5;
  -webkit-font-smoothing: antialiased;
}

a {
  color: var(--ink);
  text-decoration: none;
}

a:hover {
  color: var(--primary);
}

img {
  max-width: 100%;
  height: auto;
  display: block;
}

h1, h2, h3, h4, h5, h6 {
  color: var(--ink);
  font-weight: 400;
  line-height: 1.2;
}

h1 { font-size: 48px; letter-spacing: -1.5px; }
h2 { font-size: 36px; letter-spacing: -0.72px; }
h3 { font-size: 26px; letter-spacing: -0.325px; }
h4 { font-size: 22px; letter-spacing: -0.11px; }
h5 { font-size: 18px; font-weight: 600; }
h6 { font-size: 16px; font-weight: 600; }

p {
  color: var(--body);
  font-size: 16px;
  line-height: 1.5;
  margin-bottom: 1rem;
}

.container {
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
}

.site-nav {
  background-color: var(--canvas);
  border-bottom: 1px solid var(--hairline);
  position: sticky;
  top: 0;
  z-index: 100;
}

.nav-inner {
  display: flex;
  align-items: center;
  justify-content: space-between;
  height: 64px;
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
}

.nav-logo {
  font-size: 18px;
  font-weight: 600;
  color: var(--ink);
  letter-spacing: -0.3px;
}

.nav-logo span {
  color: var(--primary);
}

.nav-links {
  display: flex;
  align-items: center;
  gap: 32px;
  list-style: none;
}

.nav-links a {
  font-size: 14px;
  font-weight: 500;
  color: var(--body);
}

.nav-links a:hover {
  color: var(--ink);
}

.nav-hamburger {
  display: none;
  background: none;
  border: none;
  cursor: pointer;
  padding: 8px;
  flex-direction: column;
  gap: 5px;
}

.nav-hamburger span {
  display: block;
  width: 22px;
  height: 2px;
  background: var(--ink);
  border-radius: 2px;
}

.hero {
  padding: 80px 0;
  border-bottom: 1px solid var(--hairline);
}

.hero-eyebrow {
  display: inline-block;
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  background: var(--surface-strong);
  padding: 4px 10px;
  border-radius: 9999px;
  margin-bottom: 24px;
}

.hero h1 {
  font-size: 72px;
  letter-spacing: -2.16px;
  line-height: 1.1;
  color: var(--ink);
  max-width: 800px;
  margin-bottom: 24px;
}

.hero-lead {
  font-size: 18px;
  line-height: 1.6;
  color: var(--body);
  max-width: 600px;
  margin-bottom: 0;
}

.section {
  padding: 80px 0;
}

.section-label {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 16px;
}

.section-title {
  font-size: 36px;
  letter-spacing: -0.72px;
  color: var(--ink);
  margin-bottom: 48px;
}

.cards-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 24px;
}

.card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: 12px;
  overflow: hidden;
}

.card-image {
  width: 100%;
  height: 200px;
  object-fit: cover;
}

.card-body {
  padding: 24px;
}

.card-category {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 8px;
}

.card-title {
  font-size: 20px;
  font-weight: 400;
  letter-spacing: -0.2px;
  color: var(--ink);
  margin-bottom: 12px;
  line-height: 1.3;
}

.card-excerpt {
  font-size: 14px;
  color: var(--body);
  line-height: 1.5;
  margin-bottom: 16px;
}

.card-link {
  font-size: 14px;
  font-weight: 500;
  color: var(--ink);
  border-bottom: 1px solid var(--hairline-strong);
  padding-bottom: 1px;
}

.article-hero {
  padding: 80px 0 48px;
  border-bottom: 1px solid var(--hairline);
}

.article-meta {
  display: flex;
  align-items: center;
  gap: 16px;
  margin-bottom: 24px;
}

.article-category {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  background: var(--surface-strong);
  padding: 4px 10px;
  border-radius: 9999px;
}

.article-date {
  font-size: 13px;
  color: var(--muted);
}

.article-title {
  font-size: 48px;
  letter-spacing: -1.5px;
  line-height: 1.1;
  color: var(--ink);
  margin-bottom: 20px;
  max-width: 800px;
}

.article-lead {
  font-size: 18px;
  line-height: 1.6;
  color: var(--body);
  max-width: 680px;
}

.article-image-hero {
  margin-top: 48px;
  border-radius: 12px;
  width: 100%;
  max-height: 480px;
  object-fit: cover;
  border: 1px solid var(--hairline);
}

.article-content {
  padding: 64px 0;
}

.article-layout {
  display: grid;
  grid-template-columns: 1fr 300px;
  gap: 64px;
  align-items: start;
}

.article-body h2 {
  font-size: 28px;
  letter-spacing: -0.5px;
  color: var(--ink);
  margin: 48px 0 16px;
  padding-top: 48px;
  border-top: 1px solid var(--hairline);
}

.article-body h2:first-child {
  margin-top: 0;
  padding-top: 0;
  border-top: none;
}

.article-body h3 {
  font-size: 22px;
  letter-spacing: -0.2px;
  color: var(--ink);
  margin: 32px 0 12px;
  font-weight: 400;
}

.article-body p {
  font-size: 16px;
  line-height: 1.7;
  color: var(--body);
  margin-bottom: 20px;
}

.article-body ul,
.article-body ol {
  padding-left: 24px;
  margin-bottom: 20px;
}

.article-body li {
  font-size: 16px;
  line-height: 1.7;
  color: var(--body);
  margin-bottom: 6px;
}

.species-card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: 12px;
  padding: 24px;
  margin: 32px 0;
}

.species-card h3 {
  margin-top: 0 !important;
  margin-bottom: 4px !important;
  padding-top: 0 !important;
}

.species-name {
  font-size: 13px;
  font-style: italic;
  color: var(--muted);
  margin-bottom: 12px;
}

.species-detail {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 8px 16px;
  margin-top: 16px;
  font-size: 14px;
}

.species-detail dt {
  color: var(--muted);
  font-weight: 500;
}

.species-detail dd {
  color: var(--body-strong);
}

.article-body figure {
  margin: 32px 0;
}

.article-body figure img {
  width: 100%;
  border-radius: 8px;
  border: 1px solid var(--hairline);
}

.article-body figcaption {
  font-size: 13px;
  color: var(--muted);
  margin-top: 8px;
}

.sources-list {
  list-style: none;
  padding: 0 !important;
}

.sources-list li {
  padding: 12px 0;
  border-bottom: 1px solid var(--hairline-soft);
  font-size: 14px;
}

.sources-list li:last-child {
  border-bottom: none;
}

.sources-list a {
  color: var(--ink);
  text-decoration: underline;
  text-underline-offset: 3px;
}

.article-sidebar {
  position: sticky;
  top: 88px;
}

.sidebar-block {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: 12px;
  padding: 24px;
  margin-bottom: 24px;
}

.sidebar-title {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 16px;
}

.sidebar-links {
  list-style: none;
  padding: 0;
}

.sidebar-links li {
  padding: 10px 0;
  border-bottom: 1px solid var(--hairline-soft);
}

.sidebar-links li:last-child {
  border-bottom: none;
}

.sidebar-links a {
  font-size: 14px;
  color: var(--body);
  line-height: 1.4;
}

.sidebar-links a:hover {
  color: var(--ink);
}

.page-hero {
  padding: 64px 0 48px;
  border-bottom: 1px solid var(--hairline);
}

.page-title {
  font-size: 48px;
  letter-spacing: -1.5px;
  color: var(--ink);
  margin-bottom: 16px;
}

.page-lead {
  font-size: 18px;
  color: var(--body);
  max-width: 600px;
  margin-bottom: 0;
}

.page-content {
  padding: 64px 0;
  max-width: 720px;
}

.page-content h2 {
  font-size: 26px;
  letter-spacing: -0.3px;
  color: var(--ink);
  margin: 40px 0 16px;
  padding-top: 40px;
  border-top: 1px solid var(--hairline);
  font-weight: 400;
}

.page-content h2:first-child {
  margin-top: 0;
  padding-top: 0;
  border-top: none;
}

.page-content p {
  font-size: 16px;
  line-height: 1.7;
  color: var(--body);
  margin-bottom: 16px;
}

.page-content ul {
  padding-left: 20px;
  margin-bottom: 16px;
}

.page-content li {
  font-size: 16px;
  line-height: 1.7;
  color: var(--body);
  margin-bottom: 4px;
}

.contact-form {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: 12px;
  padding: 32px;
  max-width: 560px;
  margin-top: 48px;
}

.contact-form h3 {
  font-size: 22px;
  color: var(--ink);
  margin-bottom: 24px;
  font-weight: 400;
}

.form-group {
  margin-bottom: 20px;
}

.form-group label {
  display: block;
  font-size: 14px;
  font-weight: 500;
  color: var(--body-strong);
  margin-bottom: 6px;
}

.form-group input,
.form-group textarea {
  width: 100%;
  background: var(--surface-card);
  color: var(--ink);
  border: 1px solid var(--hairline-strong);
  border-radius: 8px;
  padding: 12px 16px;
  height: 44px;
  font-size: 15px;
  font-family: inherit;
  outline: none;
  transition: border-color 0.15s;
}

.form-group textarea {
  height: auto;
  resize: vertical;
  min-height: 120px;
}

.form-group input:focus,
.form-group textarea:focus {
  border-color: var(--ink);
}

.form-group input.error,
.form-group textarea.error {
  border-color: var(--error);
}

.form-error {
  font-size: 13px;
  color: var(--error);
  margin-top: 4px;
  display: none;
}

.btn-submit {
  display: inline-flex;
  align-items: center;
  background: var(--ink);
  color: var(--canvas);
  font-size: 14px;
  font-weight: 500;
  padding: 12px 20px;
  height: 44px;
  border-radius: 8px;
  border: none;
  cursor: pointer;
  font-family: inherit;
  transition: background 0.15s;
}

.btn-submit:hover {
  background: #3a3930;
}

.btn-submit:disabled {
  opacity: 0.6;
  cursor: default;
}

.form-success {
  display: none;
  background: #f0faf6;
  border: 1px solid var(--success);
  border-radius: 8px;
  padding: 16px;
  font-size: 15px;
  color: var(--success);
  margin-top: 16px;
}

.form-success.visible {
  display: block;
}

.site-footer {
  background: var(--canvas);
  border-top: 1px solid var(--hairline);
  padding: 64px 0 48px;
}

.footer-grid {
  display: grid;
  grid-template-columns: 2fr 1fr 1fr 1fr;
  gap: 48px;
  margin-bottom: 48px;
}

.footer-brand-logo {
  font-size: 18px;
  font-weight: 600;
  color: var(--ink);
  letter-spacing: -0.3px;
}

.footer-brand-logo span {
  color: var(--primary);
}

.footer-brand p {
  font-size: 14px;
  color: var(--body);
  max-width: 240px;
  margin-top: 12px;
  line-height: 1.6;
  margin-bottom: 0;
}

.footer-col-title {
  font-size: 13px;
  font-weight: 600;
  color: var(--ink);
  margin-bottom: 16px;
}

.footer-links {
  list-style: none;
  padding: 0;
}

.footer-links li {
  margin-bottom: 10px;
}

.footer-links a {
  font-size: 14px;
  color: var(--body);
}

.footer-links a:hover {
  color: var(--ink);
}

.footer-bottom {
  display: flex;
  align-items: flex-start;
  justify-content: space-between;
  gap: 32px;
  padding-top: 24px;
  border-top: 1px solid var(--hairline);
}

.footer-copyright {
  font-size: 13px;
  color: var(--muted);
  margin-bottom: 0;
  white-space: nowrap;
}

.footer-disclaimer {
  font-size: 12px;
  color: var(--muted-soft);
  max-width: 480px;
  line-height: 1.5;
  margin-bottom: 0;
}

.cookie-banner {
  position: fixed;
  bottom: 24px;
  left: 50%;
  transform: translateX(-50%);
  background: var(--ink);
  color: var(--canvas);
  border-radius: 12px;
  padding: 20px 24px;
  max-width: 600px;
  width: calc(100% - 48px);
  display: flex;
  align-items: center;
  gap: 16px;
  z-index: 9999;
}

.cookie-banner p {
  flex: 1;
  font-size: 13px;
  color: var(--canvas);
  margin: 0;
  line-height: 1.5;
}

.cookie-banner a {
  color: var(--muted-soft);
  text-decoration: underline;
}

.cookie-actions {
  display: flex;
  gap: 8px;
  flex-shrink: 0;
}

.btn-cookie-accept {
  background: var(--primary);
  color: #fff;
  border: none;
  border-radius: 8px;
  font-size: 13px;
  font-weight: 500;
  padding: 8px 16px;
  cursor: pointer;
  font-family: inherit;
  white-space: nowrap;
}

.btn-cookie-reject {
  background: transparent;
  color: var(--canvas);
  border: 1px solid rgba(255,255,255,0.25);
  border-radius: 8px;
  font-size: 13px;
  font-weight: 500;
  padding: 8px 16px;
  cursor: pointer;
  font-family: inherit;
  white-space: nowrap;
}

.last-updated {
  font-size: 13px;
  color: var(--muted);
  margin-top: 48px;
  padding-top: 24px;
  border-top: 1px solid var(--hairline-soft);
}

.breadcrumb {
  font-size: 13px;
  color: var(--muted);
  margin-bottom: 24px;
}

.breadcrumb a {
  color: var(--muted);
}

.breadcrumb span {
  margin: 0 6px;
}

@media (max-width: 1024px) {
  .hero h1 { font-size: 56px; letter-spacing: -1.5px; }
  .article-layout { grid-template-columns: 1fr; }
  .article-sidebar { position: static; }
}

@media (max-width: 768px) {
  .nav-links { display: none; }
  .nav-links.open {
    display: flex;
    flex-direction: column;
    position: absolute;
    top: 64px;
    left: 0;
    right: 0;
    background: var(--canvas);
    border-bottom: 1px solid var(--hairline);
    padding: 8px 24px;
    gap: 0;
  }
  .nav-links.open li {
    border-bottom: 1px solid var(--hairline-soft);
  }
  .nav-links.open a {
    display: block;
    padding: 14px 0;
  }
  .nav-hamburger { display: flex; }
  .site-nav { position: relative; }

  .hero { padding: 48px 0; }
  .hero h1 { font-size: 36px; letter-spacing: -1px; }
  .hero-lead { font-size: 16px; }

  .section { padding: 48px 0; }
  .cards-grid { grid-template-columns: 1fr; }

  .article-title { font-size: 32px; letter-spacing: -0.8px; }
  .article-hero { padding: 48px 0 32px; }

  .footer-grid { grid-template-columns: 1fr 1fr; gap: 32px; }

  .cookie-banner { flex-direction: column; align-items: flex-start; }
  .cookie-actions { width: 100%; }
  .btn-cookie-accept, .btn-cookie-reject { flex: 1; text-align: center; }
}

@media (max-width: 480px) {
  .footer-grid { grid-template-columns: 1fr; }
  .species-detail { grid-template-columns: 1fr; }
  .page-title { font-size: 32px; }
  .contact-form { padding: 24px; }
}